Reference citation: Kottelat, M. and J. Freyhof. 2007. Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp.
Source citation: Froese, R. and D. Pauly. Editors. 2018. FishBase. World Wide Web electronic publication. www.fishbase.org, version (10/2018).